Bernard Munyagishari is accused of being a militia leader who orchestrated the murder of Tutsis and moderate Hutus Gisenyi in 1994.
He was caught in an operation mounted by the Congolese army in collaboration with a team of tracking by the Tribunal of the United Nations tent of genocide suspects.
Some 800,000 ethnic Tutsis and moderate Hutus died in the 100-day genocide.
Nine other Rwandan, accused of being key authors of the genocide are still at large.
The Church attacksCorrespondent of the BBC Will Ross Africa, explains Mr. Munyagishari, once a teacher and football referee, was apprehended in North Kivu in Eastern Congo DR - where many of those who participated in the 1994 genocide were hiding and causing the suffering of the Congolese people.
Mr. Munyagishari is facing charges of genocide, murder and rape and is should be provided to the Tribunal International Criminal for Rwanda (ICTR), the Court in Tanzania established to try the ringleaders of the genocide.
According to the indictment, he was the head of the militia Interahamwe Hutu in Gisenyi in the West of the country.
He trained fighters in forests in Rwanda and their army to wipe out the ethnic Tutsi population.
He is also accused of inciting attacks on churches where people had sought refuge.
The indictment said he ordered a group of female militiamen, led by his wife, in sexually tortured Tutsi women before killing.
Our correspondent says that one of the nine suspects of genocide in the large is Felicien Kabuga, that many believe have been protected for years by powerful politicians in the Kenya.
Last week, the Chief Prosecutor of the ICTR court stated that the Kenyan Government could more be done to apprehend him.
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